Despite the anagram, the main protagonists are not witches. Like the comics, the series follows five ordinary girls: Will, Irma, Taranee, Cornelia and Hay Lin who become endowed with magical powers and become the "Guardians of the Veil".
They must save Meridian from the evil sorcerer Phobos and Cedric who are searching for Phobos' sister, the long lost princess of Meridian. It is later revealed the princess is Cornelia's best friend Elyon, and the Guardians then set about saving her from Phobos. When Meridian is freed from evil and Elyon takes the throne, a new mysterious sorceress named Nerissa frees Phobos's top henchmen and reforms them as the Knights of Vengeance. Once the Guardians learn more about the sorceress and her evil plan of reuniting former Guardians, they are able to defeat the Knights only to have more powerful Knights of Destruction, plus the old Guardians attack them.
The Guardians chief ally is Hay Lin's grandmother Yan Lin, a former Guardian herself, and the one that taught the girls about their magical destiny. They are also helped by Caleb, a heroic soldier from Meridian, leader of the rebellion against Phobos, and Blunk, a frog-like goblin creature (known as a Passling) who takes things from the human world to Meridian (and vice-versa), humorously mistaking everyday objects for other things or items of value. Matt, Will's boyfriend, accidentally learns about Meridian and when he sees all the trouble on it going on, he learns how to become a warrior to help them. They are also helped by the Oracle, leader of the Universe in Kandrakar, who was the one who chose the Guardians.
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